Mishana Tyrannulet forages actively in the canopy and subcanopy, typically making short flights between perches and occasionally flicking wings slightly outward. The posture is relatively horizontal for a flycatcher, as is typical of the genus Zimmerius. The tail usually is held angled downward, and occasionally is flicked upward (Alvarez and Whitney 2001).
Little information is available on territoriality in Mishana Tyrannulet. Individuals often can be found in the same location (i.e. tree or groups of trees) over fairly long time periods (months at least), suggesting that this species maintains stable home ranges (Alvarez and Whitney 2001). Individuals regularly approach and vocalize in response to playback in a manner typical of type A territorial species.
Undescribed; Mishana Tyrannulet probably is at least socially monogamous.
Social and interspecific behavior
Little information is available on social and interspecific behavior of Mishana Tyrannulet. The species does not typically join mixed flocks, but appears to move in pairs at least some of the time. Alvarez and Whitney (2001) report that an individual foraging on mistletoe chased away an Ochre-bellied Flycatcher (Mionectes oleagineus) but later tolerated the presence of a conspecific.
No predation events have been documented, but Alvarez and Whitney (2001) report that the species responded behaviorally to imitations of Ferruginous Pygmy-Owl (Glaucidium brasilianum) whistles.