This flycatcher is endemic to Robinson Crusoe Island, in the Juan Fernandez Archipelago of Chile. It is found in the mid story of various wooded habitats, although the preference is for native forest, such as Luma (Nothomyrica fernandeziana). This tit-tyrant is larger than its close relatives on the mainland, and it forages in a slow and deliberate manner quite unlike the frenetic hyperactivity of smaller Anairetes. It may sit still for stretches at a time, without much activity. Little is known about its vocal behavior but preliminary observations show that the male and female perform a duet, and that the voice is quite dissimilar to its closest relative (based on molecular data), the Tufted Tit-Tyrant (Anairetes parulus). The population was estimated to be 5000 individuals and stable in the 80s; it is listed as Near Threatened by the IUCN.