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Guanay Cormorant Phalacrocorax bougainvillii

  • Order: Suliformes
  • Family: Phalacrocoracidae
  • Monotypic
  • Authors: Mark A. Baran and Michael G. Harvey


Barbraud, C., A. Bertrand, M. Bouchón, A. Chaigneau, K. Delord, H. Demarcq, O. Gimenez, M. Gutiérrez Torero, D. Gutiérrez, R. Oliveros-Ramos, G. Passuni, Y. Tremblay, and S. Bertrand. 2017. Density dependence, prey accessibility and prey depletion by fisheries drive Peruvian seabird population dynamics. Ecography: in press.

Bertellotti, M., J. A. Donázar, G. Blanco, and M. G. Forero. 2003. Imminent extinction of the Guanay Cormorant on the Atlantic South American coast: a conservation concern? Biodiversity and Conservation 12: 743–747.

BirdLife International. 2010. Species factsheet: Guanay Cormorant Phalacrocorax bougainvillii. Downloaded from BirdLife International on 19 May 2010.

Blake, E. R. 1977. Manual of neotropical birds. Volume 1. University of Chicago Press, Chicago, Illinois.

Devillers, P., and J. A. Terchuren. 1978. Relationships between the blue-eyed shags of South America. Gerfaut 68: 53-86.

Duffy, D.C. 1980. Patterns of piracy by Peruvian seabirds: A depth hypothesis. Ibis 122: 521-525.

Duffy, D. C. 1983a. Competition for nesting space among Peruvian guano birds. Auk 100: 680-688.

Duffy, D. C. 1983b. Environmental uncertainty and commercial fishing: effects on Peruvian guano birds. Biological Conservation 26: 227-238.

Duffy, D. C. 1983c. The foraging ecology of Peruvian seabirds. Auk 100: 800-810.

Duffy, D.C. 1983d. The ecology of tick parasitism on densely nesting Peruvian seabirds. Ecology 64: 110-119.

Hilty, S. L., and W. L. Brown. 1986. A guide to the birds of Colombia. Princeton University Press, Princeton, New Jersey.

Jaramillo, A. 2003. Birds of Chile. Princeton University Press, Princeton, New Jersey.

Johnson, A. W. 1965. The birds of Chile and adjacent regions of Argentina, Bolivia and Peru. Volume I. Platt Establecimientos Gráficos, Buenos Aires.

Jordan, R. 1966. The predation of guano birds on the Peruvian anchovy. California Cooperative Oceanic Fisheries Investigations 11: 105-109.

Kennedy, M., R. D. Gray, and H. G. Spencer. 2000. The phylogenetic relationships of the shags and cormorants: can sequence data resolve a disagreement between behavior and morphology? Molecular Phylogenetics and Evolution 17: 345-359.

Murphy R. C. 1936. Oceanic birds of South America. American Museum of Natural History, New York, New York.

Ridgely, R. S., and P. J. Greenfield. 2001. The birds of Ecuador: status, distribution, and taxonomy. Cornell University Press, Ithaca, New York.

Ridgely, R. S., and J. A. Gwynne. 1989. A guide to the birds of Panama with Costa Rica, Nicaragua, and Honduras. Princeton University Press, Princeton, New Jersey.

Sepulveda, M. S ., R. L. Palma, and H. Ochoa-Acuña. 1997. New records of feather lice from some seabirds in Chile. Journal of Wildlife Diseases 33: 371-372.

Siegel-Causey, D. 1988. Phylogeny of the Phalacrocoracidae. Condor 90: 885-905.

Weimerskirch, H., S. Bertrand, J. Silva, J. C. Marques, and E. Goya. 2010. Use of social information in seabirds: compass rafts indicate the heading of food patches. PLoS ONE 5(3): e9928.

Zavalaga, C. B., and R. Paredes. 1999. Foraging behaviour and diet of the Guanay Cormorant. South African Journal of Marine Science 21: 251-258.

Recommended Citation

Baran, M. A. and M. G. Harvey (2011). Guanay Cormorant (Phalacrocorax bougainvillii), version 1.0. In Neotropical Birds Online (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA.